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What's in a name?
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As was the case for Juliet, for many of us the name of a particular plant in the garden makes little difference. We enjoy the look, smell, feel of a plant and that is enough. From that perspective, it is hard to imagine the obsession of the Swedish scientist, Carl Linnaeus, (1707-1778), who spend much of his life classifying and naming plants and animals. By 1753, Linnaeus had published the two volume Species Plantarum in which he carefully documented some 6000 species of plants. While I can not imagine spending my days characterizing plants, the Linnaean system is most useful.
Linnaeus grouped plants within a Kingdom, Vegetabilia, and divided that kingdom into classes, orders, genera and species. Within this heirarchy, Linnaeus gave long descriptive names to each plant. For instance, the tomato was described as Solanum caule inermi herbaceo foliis pinnatis incisis, racemis simplicibus meaning this plant belonged to the nightshade genus (Solanum) and was characterized by an having an herbaceous spineless stem, pinnately incised leaves, and simple racemes (flowers with short stalks). As you might imagine, Linnaeus soon replaced this cumbersome naming system with short binomial names or nomina trivialia. The tomato became Solanum lycopersicum (lyco - wolf; persicum - peach-like). (While it is known that the Brazilian wolf apple, Solanum lycocarpum, are eaten by Maned wolves, it is not know if this is the origin of the wolf reference.1) This simple naming system, with italicized genus name first (first letter capitalized) and the species name second (all lowercase), remains a standard today.
Now if you've been to a nursery lately, you will see dozens of varieties of tomatoes like 'Big Boy' or 'Beafsteak'. These are appended to the name, e.g. S. lycopersicum 'Red Brandywine'. In places like Ecuador and Chili, there are naturally occurring variations like S. lycopersicum var. cerasiforme. Of course there are also many relatives of the tomato like our eggplant, Solanum melongena L. (The L. at the end denotes that Linnaeus named this plant - you may see other letters or names referencing the authority who identified the plant appended to a name.) I should mention that although, Linnaeus had named the tomato in 1753, another authority renamed it in 1768 as Lycopersicon esculentum Miller (this name means edible wolf-peach). It is Miller's name for the tomato that has actually been in use in the scientific literature for some 200 years. It is only recently that genetic research reshuffled the plant back into the nightshade genus.
No wonder we use the Nahuatl name (old Aztec), tomato.
Although learning the binomial names for plants can be a challenge, it can also help you understand relationships between plants and help you predict cultural requirements as well as disease and pest susceptibility. It can also prevent confusion among common names. At our recent plant sale, we debated how to organize plants. Should sea holly be sorted under S for sea, H for holly or E for Eryngium? In some instances, we had plants under a variety of letters to help people locate them. For species with many common names like our native Nyssa sylvatica (e.g. Sour Gum, Black Gum, Black Tupelo, Pepperidge), knowing the scientific name can be quite helpful.
I will end this note with an apology to whoever bought the Waldsteina or barren strawberry I offered at the plant sale. While I purchased the plant believing it was our native W. fragarioides, it is now in bloom in my garden and is almost certainly the more attractive European W. ternata. I see this often in plants marketed as native and it is a problem from the prospective of maintaining biodiversity and preserving local gene pools. While the appearance of plants may be quite similar, subtle differences in genetics give the plants quite different properties within an ecosystem. For instance, many cousins in the tomato family bear fruit, but they are not safe to eat because of their high solanum content. Ah well, W. ternata makes a lovely ground cover and I am certainly enjoying its cheery yellow blooms.
May toads nestle in your flower beds and help you tend your beauties this spring.
Resources and Interesting Tidbits
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Some taxonomy basics: The basic categories for classifying plants are: Domain, Kingdom, Phylum, Class, Order, Family, Tribe, Genus, Section, Series, Species, Variety (or Cultivar). The physical features of plants become more and more similar as one proceeds to finer levels of classification.One of many pneumonics to remember this particular organizational structure is: Daring Kangaroos Play Cellos, Orangutans Fiddle, The Gorillas Sings Special Songs. (Variants on the primary seven groups include: "Karen, please come over for great sex;" "King Philip came over for great sex;" etc.) Linnaeus originally defined three Kingdoms (including Mineralia for minerals) which have evolved over time as our understanding of genetics have improved, (see Phylogenomics). Here is a helpful table showing the historical progression of kingdoms:
Plant Divisions or Phyla: Phylum names traditionally end in phyta. As with kingdoms, these divisions or phyla are evolving and the members shifting as our knowledge and genetic techniques improve. The initial four divisions: Thallophyta (bacteria, algae, fungi), Bryophyta (mosses and liverworts, Pteridophyta (ferns and cubmosses) and Spermatophyta (seed plants: gymnosperms and angiosperms) have been split and expanded and there is debate about these divisions. The groupings are largely based on reproductive characteristics and tissue structure. This table below is from the Wiki on Phylum (see Wiki Ansers on Plant phylum or UCMP Berkeley for a different scheme: http://www.ucmp.berkeley.edu/help/index/plantae.html)
Classes The division Ginkgophyta is by far the simplest as there are so few members (There are seven principle genera: Baiera†, Ginkgo, Ginkgoites†, Ginkgoïdium†, Phoenicpsis†, Polyspermophyllum†, Trichopitys† with only one living species). Thus the division has one class: Ginkgoopsida (classes typically end in -sida); one order: Ginkgoales (orders typically end in -ales) and one family: Ginkgoaceae (families typically end in -ceae). There is some debate however that the order Ginkgoales should be placed in a different class. As ginkgoes have reproductive structures similar to that of the cycads that could be classed within Cycadophyta. Alternatively they share vegetative morphology of the conifers and could be classed within Pinophyta. The solution was to give them a class unto themselves, for now, (see UCMP Systematics http://www.ucmp.berkeley.edu/seedplants/ginkgoales/ginkgosy.html ) In the largest complex phylum, Anthrophyta previously called Magnoliophyta (flowering plants), classification becomes even more contentious. The flowering plants have historically been divided into two Classes based on the number of cotyledons on the embryo. Those flowering plants with two cotyledons are called dicots while those with only one are called monocots (see UCMP Berkeley Discussion of Classes). Systematists argue for more classes, different classes and more levels. For simplicity sake, there are two classes: Within classes, are Orders which organize finer detail. It is a subjective art choosing which characteristics to consider forming into a groups. While there is much debate on this organization, here is one useful classification structure.
"The APG II system does not assign the monocots to a taxonomic rank, instead recognizing a monocots clade. This system recognizes ten orders of monocots and two families of monocots not yet assigned to any order" (Wiki Monocots see discussion of Cladistics) Family
Genus The differences between members of a genus become more subtle. They will share similarities in physical characteristics that belie their common genetics. However, it is sometimes discovered that the physical characteristics are coincidental, in which case members will be shifted and to other groupings. For instance, the goldenrods were once lumped together within the genus Solidago. It was recently found that the Grass-leaved Goldenrods were not that closely related and have been moved to the genus Euthamia (see for instance John Semple Goldenrods). Similarity of flowers and fruits is the most widely used feature, although roots, stems, buds, and leaves are also used. Common names typically apply to genera. For example Hemerocallis is the genus of daylilies, Acer is the genus of maples, and Viburnum the genus of viburnum. (Note when a genus name is used as a common name, it is not capitalized). Species Species are a group of plants that conform to
certain fixed properties. Naturally occurring populations interbreed at
maturity and sustain these properties. While there may be clusters of
variations or phenotypes, they are sufficiently similar to be grouped
together. Modern genetic techniques have been helpful to define particular
genotypes associated with specific species. (See Wiki on
Species
and Species Problem).
Species naming conventions Common names are lower case unless the word is derived from a surname or proper noun, e.g. tomato, Kentucky coffee tree, black-eyed Susan, Douglas fir, etc. Botanical names are often used as common names in which case they are set in lower case and not italicized, e.g. rhododendron, hosta, iris, etc. Botanical names for a given species are given as genus name or abbreviation followed by species name and variety name as follows.
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